1 c2pn5A_
100.0
14
PDB header: immune systemChain: A: PDB Molecule: thioester-containing protein i;
PDBTitle: crystal structure of tep1r
2 c3cu7A_
100.0
14
PDB header: immune systemChain: A: PDB Molecule: complement c5;
PDBTitle: human complement component 5
3 c2b39B_
100.0
16
PDB header: immune systemChain: B: PDB Molecule: c3;
PDBTitle: structure of mammalian c3 with an intact thioester at 3a resolution
4 c2a73B_
100.0
15
PDB header: immune systemChain: B: PDB Molecule: complement c3;
PDBTitle: human complement component c3
5 c3g6jB_
100.0
15
PDB header: immune systemChain: B: PDB Molecule: complement c3 alpha chain;
PDBTitle: c3b in complex with a c3b specific fab
6 c3pvmB_
100.0
18
PDB header: immune systemChain: B: PDB Molecule: cobra venom factor;
PDBTitle: structure of complement c5 in complex with cvf
7 d1hzfa_
100.0
19
Fold: alpha/alpha toroidSuperfamily: Terpenoid cyclases/Protein prenyltransferasesFamily: Complement components
8 c1hzfA_
100.0
19
PDB header: immune systemChain: A: PDB Molecule: complement factor c4a;
PDBTitle: c4adg fragment of human complement factor c4a
9 d1c3da_
100.0
19
Fold: alpha/alpha toroidSuperfamily: Terpenoid cyclases/Protein prenyltransferasesFamily: Complement components
10 c3hrzA_
100.0
13
PDB header: immune systemChain: A: PDB Molecule: cobra venom factor;
PDBTitle: cobra venom factor (cvf) in complex with human factor b
11 d1qqfa_
100.0
16
Fold: alpha/alpha toroidSuperfamily: Terpenoid cyclases/Protein prenyltransferasesFamily: Complement components
12 c2qkiA_
100.0
14
PDB header: immune system/hydrolase inhibitorChain: A: PDB Molecule: complement c3;
PDBTitle: human c3c in complex with the inhibitor compstatin
13 c3hs0B_
100.0
14
PDB header: immune systemChain: B: PDB Molecule: cobra venom factor;
PDBTitle: cobra venom factor (cvf) in complex with human factor b
14 c2a74B_
99.9
11
PDB header: immune systemChain: B: PDB Molecule: complement component c3c;
PDBTitle: human complement component c3c
15 c2p9rA_
99.7
23
PDB header: signaling proteinChain: A: PDB Molecule: alpha-2-macroglobulin;
PDBTitle: human alpha2-macroglogulin is composed of multiple domains,2 as predicted by homology with complement component c3
16 d2sqca1
99.6
16
Fold: alpha/alpha toroidSuperfamily: Terpenoid cyclases/Protein prenyltransferasesFamily: Terpene synthases
17 c3hrzC_
99.6
10
PDB header: immune systemChain: C: PDB Molecule: cobra venom factor;
PDBTitle: cobra venom factor (cvf) in complex with human factor b
18 c1o79A_
99.6
20
PDB header: isomeraseChain: A: PDB Molecule: squalene--hopene cyclase;
PDBTitle: structures of human oxidosqualene cyclase inhibitors bound2 to a homolgous enzyme
19 c1w6kA_
99.4
18
PDB header: isomeraseChain: A: PDB Molecule: lanosterol synthase;
PDBTitle: structure of human osc in complex with lanosterol
20 d1edya_
99.4
15
Fold: Common fold of diphtheria toxin/transcription factors/cytochrome fSuperfamily: Alpha-macroglobulin receptor domainFamily: Alpha-macroglobulin receptor domain
21 d1ayoa_
not modelled
99.4
16
Fold: Common fold of diphtheria toxin/transcription factors/cytochrome fSuperfamily: Alpha-macroglobulin receptor domainFamily: Alpha-macroglobulin receptor domain
22 d1bv8a_
not modelled
99.4
14
Fold: Common fold of diphtheria toxin/transcription factors/cytochrome fSuperfamily: Alpha-macroglobulin receptor domainFamily: Alpha-macroglobulin receptor domain
23 d3es6b1
not modelled
99.4
11
Fold: Immunoglobulin-like beta-sandwichSuperfamily: E set domainsFamily: SVA-like
24 d2sqca2
not modelled
99.3
15
Fold: alpha/alpha toroidSuperfamily: Terpenoid cyclases/Protein prenyltransferasesFamily: Terpene synthases
25 d1w6ka1
not modelled
99.3
18
Fold: alpha/alpha toroidSuperfamily: Terpenoid cyclases/Protein prenyltransferasesFamily: Terpene synthases
26 c2qkiC_
not modelled
99.2
7
PDB header: immune system/hydrolase inhibitorChain: C: PDB Molecule: complement c3;
PDBTitle: human c3c in complex with the inhibitor compstatin
27 d3dssb1
not modelled
98.6
17
Fold: alpha/alpha toroidSuperfamily: Terpenoid cyclases/Protein prenyltransferasesFamily: Protein prenyltransferases
28 c3e37B_
not modelled
98.2
13
PDB header: transferaseChain: B: PDB Molecule: protein farnesyltransferase subunit beta;
PDBTitle: protein farnesyltransferase complexed with bisubstrate2 ethylenediamine scaffold inhibitor 5
29 d2h6fb1
not modelled
98.1
13
Fold: alpha/alpha toroidSuperfamily: Terpenoid cyclases/Protein prenyltransferasesFamily: Protein prenyltransferases
30 c1cwvA_
not modelled
97.8
10
PDB header: structural proteinChain: A: PDB Molecule: invasin;
PDBTitle: crystal structure of invasin: a bacterial integrin-binding protein
31 c3q7aB_
not modelled
97.7
14
PDB header: transferase/transferase inhibitorChain: B: PDB Molecule: farnesyltransferase beta subunit;
PDBTitle: cryptococcus neoformans protein farnesyltransferase in complex with2 fpp and l-778,123
32 c2pmvB_
not modelled
97.4
11
PDB header: transport proteinChain: B: PDB Molecule: gastric intrinsic factor;
PDBTitle: crystal structure of human intrinsic factor- cobalamin complex at 2.62 a resolution
33 c1tnuL_
not modelled
97.4
12
PDB header: transferaseChain: L: PDB Molecule: geranylgeranyl transferase type i beta subunit;
PDBTitle: rat protein geranylgeranyltransferase type-i complexed with2 a ggpp analog and a gcincckvl peptide derived from rhob
34 d1n4qb_
not modelled
97.4
12
Fold: alpha/alpha toroidSuperfamily: Terpenoid cyclases/Protein prenyltransferasesFamily: Protein prenyltransferases
35 d1w6ka2
not modelled
97.1
19
Fold: alpha/alpha toroidSuperfamily: Terpenoid cyclases/Protein prenyltransferasesFamily: Terpene synthases
36 c3draB_
not modelled
97.0
10
PDB header: transferaseChain: B: PDB Molecule: geranylgeranyltransferase type i beta subunit;
PDBTitle: candida albicans protein geranylgeranyltransferase-i2 complexed with ggpp
37 d1w8oa1
not modelled
96.8
17
Fold: Immunoglobulin-like beta-sandwichSuperfamily: E set domainsFamily: E-set domains of sugar-utilizing enzymes
38 c1l9mB_
not modelled
96.6
15
PDB header: transferaseChain: B: PDB Molecule: protein-glutamine glutamyltransferase e3;
PDBTitle: three-dimensional structure of the human transglutaminase 32 enzyme: binding of calcium ions change structure for3 activation
39 d1r76a_
not modelled
96.5
23
Fold: alpha/alpha toroidSuperfamily: Family 10 polysaccharide lyaseFamily: Family 10 polysaccharide lyase
40 d1gxma_
not modelled
96.5
23
Fold: alpha/alpha toroidSuperfamily: Family 10 polysaccharide lyaseFamily: Family 10 polysaccharide lyase
41 c2v3nA_
not modelled
96.1
13
PDB header: transport proteinChain: A: PDB Molecule: transcobalamin-2;
PDBTitle: crystallographic analysis of upper axial ligand2 substitutions in cobalamin bound to transcobalamin
42 d2afaa1
not modelled
95.1
17
Fold: alpha/alpha toroidSuperfamily: Six-hairpin glycosidasesFamily: N-acylglucosamine (NAG) epimerase
43 c2kl6A_
not modelled
94.6
19
PDB header: structural genomics, unknown functionChain: A: PDB Molecule: uncharacterized protein;
PDBTitle: solution nmr structure of the cardb domain of pf1109 from2 pyrococcus furiosus. northeast structural genomics3 consortium target pfr193a
44 c3e8vA_
not modelled
93.1
21
PDB header: structural genomics, unknown functionChain: A: PDB Molecule: possible transglutaminase-family protein;
PDBTitle: crystal structure of a possible transglutaminase-family2 protein proteolytic fragment from bacteroides fragilis
45 d1cwva1
not modelled
92.5
11
Fold: Immunoglobulin-like beta-sandwichSuperfamily: Invasin/intimin cell-adhesion fragmentsFamily: Invasin/intimin cell-adhesion fragments
46 c1wlhA_
not modelled
91.7
12
PDB header: structural proteinChain: A: PDB Molecule: gelation factor;
PDBTitle: molecular structure of the rod domain of dictyostelium2 filamin
47 d1h8la1
not modelled
91.5
13
Fold: Prealbumin-likeSuperfamily: Carboxypeptidase regulatory domain-likeFamily: Carboxypeptidase regulatory domain
48 c3irpX_
not modelled
90.5
36
PDB header: cell adhesionChain: X: PDB Molecule: uro-adherence factor a;
PDBTitle: crystal structure of functional region of uafa from staphylococcus2 saprophyticus at 1.50 angstrom resolution
49 c3kptA_
not modelled
89.9
18
PDB header: cell adhesionChain: A: PDB Molecule: collagen adhesion protein;
PDBTitle: crystal structure of bcpa, the major pilin subunit of2 bacillus cereus
50 c2h0eA_
not modelled
89.4
12
PDB header: hydrolaseChain: A: PDB Molecule: transthyretin-like protein pucm;
PDBTitle: crystal structure of pucm in the absence of substrate
51 d1qfha1
not modelled
89.3
18
Fold: Immunoglobulin-like beta-sandwichSuperfamily: E set domainsFamily: Filamin repeat (rod domain)
52 d1f00i1
not modelled
88.9
10
Fold: Immunoglobulin-like beta-sandwichSuperfamily: Invasin/intimin cell-adhesion fragmentsFamily: Invasin/intimin cell-adhesion fragments
53 c1kv3F_
not modelled
88.8
9
PDB header: transferaseChain: F: PDB Molecule: protein-glutamine gamma-glutamyltransferase;
PDBTitle: human tissue transglutaminase in gdp bound form
54 c2l0dA_
not modelled
88.8
14
PDB header: cell adhesionChain: A: PDB Molecule: cell surface protein;
PDBTitle: solution nmr structure of putative cell surface protein ma_4588 (272-2 376 domain) from methanosarcina acetivorans, northeast structural3 genomics consortium target mvr254a
55 c3qvaB_
not modelled
87.5
33
PDB header: hydrolaseChain: B: PDB Molecule: transthyretin-like protein;
PDBTitle: structure of klebsiella pneumoniae 5-hydroxyisourate hydrolase
56 c1g0dA_
not modelled
86.6
15
PDB header: transferaseChain: A: PDB Molecule: protein-glutamine gamma-glutamyltransferase;
PDBTitle: crystal structure of red sea bream transglutaminase
57 d1cwva2
not modelled
86.3
8
Fold: Immunoglobulin-like beta-sandwichSuperfamily: Invasin/intimin cell-adhesion fragmentsFamily: Invasin/intimin cell-adhesion fragments
58 c2pz4A_
not modelled
85.2
16
PDB header: cell adhesionChain: A: PDB Molecule: protein gbs052;
PDBTitle: crystal structure of spab (gbs52), the minor pilin in gram-positive2 pathogen streptococcus agalactiae
59 d1kgia_
not modelled
85.0
11
Fold: Prealbumin-likeSuperfamily: Transthyretin (synonym: prealbumin)Family: Transthyretin (synonym: prealbumin)
60 d2q3za3
not modelled
84.8
8
Fold: Immunoglobulin-like beta-sandwichSuperfamily: Transglutaminase, two C-terminal domainsFamily: Transglutaminase, two C-terminal domains
61 c1h8lA_
not modelled
84.8
13
PDB header: carboxypeptidaseChain: A: PDB Molecule: carboxypeptidase gp180 residues 503-882;
PDBTitle: duck carboxypeptidase d domain ii in complex with gemsa
62 d1uwya1
not modelled
84.6
13
Fold: Prealbumin-likeSuperfamily: Carboxypeptidase regulatory domain-likeFamily: Carboxypeptidase regulatory domain
63 d2diba1
not modelled
84.4
17
Fold: Immunoglobulin-like beta-sandwichSuperfamily: E set domainsFamily: Filamin repeat (rod domain)
64 c2r39A_
not modelled
83.9
17
PDB header: structural genomics, unknown functionChain: A: PDB Molecule: fixg-related protein;
PDBTitle: crystal structure of fixg-related protein from vibrio parahaemolyticus
65 c3mn8A_
not modelled
83.4
10
PDB header: hydrolaseChain: A: PDB Molecule: lp15968p;
PDBTitle: structure of drosophila melanogaster carboxypeptidase d isoform 1b2 short
66 c1uwyA_
not modelled
82.9
11
PDB header: hydrolaseChain: A: PDB Molecule: carboxypeptidase m;
PDBTitle: crystal structure of human carboxypeptidase m
67 c2h1xB_
not modelled
82.7
18
PDB header: hydrolaseChain: B: PDB Molecule: 5-hydroxyisourate hydrolase (formerly known as
PDBTitle: crystal structure of 5-hydroxyisourate hydrolase (formerly2 known as trp, transthyretin related protein)
68 d1vjja3
not modelled
82.6
18
Fold: Immunoglobulin-like beta-sandwichSuperfamily: Transglutaminase, two C-terminal domainsFamily: Transglutaminase, two C-terminal domains
69 d1f86a_
not modelled
82.6
13
Fold: Prealbumin-likeSuperfamily: Transthyretin (synonym: prealbumin)Family: Transthyretin (synonym: prealbumin)
70 d2e9ia1
not modelled
82.3
14
Fold: Immunoglobulin-like beta-sandwichSuperfamily: E set domainsFamily: Filamin repeat (rod domain)
71 c1f13A_
not modelled
81.7
10
PDB header: coagulation factorChain: A: PDB Molecule: cellular coagulation factor xiii zymogen;
PDBTitle: recombinant human cellular coagulation factor xiii
72 c3saeA_
not modelled
81.7
9
PDB header: lyaseChain: A: PDB Molecule: alpha-bisabolene synthase;
PDBTitle: structure of a three-domain sesquiterpene synthase: a prospective2 target for advanced biofuels production
73 d1tfpa_
not modelled
81.3
16
Fold: Prealbumin-likeSuperfamily: Transthyretin (synonym: prealbumin)Family: Transthyretin (synonym: prealbumin)
74 d1ttaa_
not modelled
80.8
11
Fold: Prealbumin-likeSuperfamily: Transthyretin (synonym: prealbumin)Family: Transthyretin (synonym: prealbumin)
75 d1g0da3
not modelled
80.7
13
Fold: Immunoglobulin-like beta-sandwichSuperfamily: Transglutaminase, two C-terminal domainsFamily: Transglutaminase, two C-terminal domains
76 d1oo2a_
not modelled
80.6
18
Fold: Prealbumin-likeSuperfamily: Transthyretin (synonym: prealbumin)Family: Transthyretin (synonym: prealbumin)
77 c2k7pA_
not modelled
80.4
14
PDB header: structural proteinChain: A: PDB Molecule: filamin-a;
PDBTitle: filamin a ig-like domains 16-17
78 c2gpzC_
not modelled
80.2
24
PDB header: hydrolaseChain: C: PDB Molecule: transthyretin-like protein;
PDBTitle: transthyretin-like protein from salmonella dublin
79 d1vjja2
not modelled
79.8
21
Fold: Immunoglobulin-like beta-sandwichSuperfamily: Transglutaminase, two C-terminal domainsFamily: Transglutaminase, two C-terminal domains
80 d1ex0a2
not modelled
77.5
11
Fold: Immunoglobulin-like beta-sandwichSuperfamily: Transglutaminase, two C-terminal domainsFamily: Transglutaminase, two C-terminal domains
81 d2q3za2
not modelled
77.4
12
Fold: Immunoglobulin-like beta-sandwichSuperfamily: Transglutaminase, two C-terminal domainsFamily: Transglutaminase, two C-terminal domains
82 d2d7ma1
not modelled
77.0
18
Fold: Immunoglobulin-like beta-sandwichSuperfamily: E set domainsFamily: Filamin repeat (rod domain)
83 d1qfha2
not modelled
76.6
10
Fold: Immunoglobulin-like beta-sandwichSuperfamily: E set domainsFamily: Filamin repeat (rod domain)
84 d1ex0a3
not modelled
76.2
17
Fold: Immunoglobulin-like beta-sandwichSuperfamily: Transglutaminase, two C-terminal domainsFamily: Transglutaminase, two C-terminal domains
85 c2xzzA_
not modelled
74.1
14
PDB header: transferaseChain: A: PDB Molecule: protein-glutamine gamma-glutamyltransferase k;
PDBTitle: crystal structure of the human transglutaminase 1 beta-barrel domain
86 c2nsmA_
not modelled
73.5
20
PDB header: hydrolaseChain: A: PDB Molecule: carboxypeptidase n catalytic chain;
PDBTitle: crystal structure of the human carboxypeptidase n (kininase i)2 catalytic domain
87 d2dmca1
not modelled
71.9
12
Fold: Immunoglobulin-like beta-sandwichSuperfamily: E set domainsFamily: Filamin repeat (rod domain)
88 c2ww8A_
not modelled
71.8
24
PDB header: cell adhesionChain: A: PDB Molecule: cell wall surface anchor family protein;
PDBTitle: structure of the pilus adhesin (rrga) from streptococcus2 pneumoniae
89 c2e9jA_
not modelled
71.7
21
PDB header: structural proteinChain: A: PDB Molecule: filamin-b;
PDBTitle: solution structure of the 14th filamin domain from human2 filamin-b
90 d2dmba1
not modelled
70.9
25
Fold: Immunoglobulin-like beta-sandwichSuperfamily: E set domainsFamily: Filamin repeat (rod domain)
91 c2k7qA_
not modelled
69.3
13
PDB header: structural proteinChain: A: PDB Molecule: filamin-a;
PDBTitle: filamin a ig-like domains 18-19
92 d1cwva3
not modelled
67.9
8
Fold: Immunoglobulin-like beta-sandwichSuperfamily: Invasin/intimin cell-adhesion fragmentsFamily: Invasin/intimin cell-adhesion fragments
93 d1v05a_
not modelled
67.4
17
Fold: Immunoglobulin-like beta-sandwichSuperfamily: E set domainsFamily: Filamin repeat (rod domain)
94 d1g0da2
not modelled
67.0
9
Fold: Immunoglobulin-like beta-sandwichSuperfamily: Transglutaminase, two C-terminal domainsFamily: Transglutaminase, two C-terminal domains
95 c3k11A_
not modelled
67.0
13
PDB header: hydrolaseChain: A: PDB Molecule: putative glycosyl hydrolase;
PDBTitle: crystal structure of putative glycosyl hydrolase (np_813087.1) from2 bacteroides thetaiotaomicron vpi-5482 at 1.80 a resolution
96 c1ksrA_
not modelled
65.8
15
PDB header: actin binding proteinChain: A: PDB Molecule: gelation factor;
PDBTitle: the repeating segments of the f-actin cross-linking2 gelation factor (abp-120) have an immunoglobulin fold, nmr,3 20 structures
97 c1qfhB_
not modelled
65.1
13
PDB header: actin binding proteinChain: B: PDB Molecule: protein (gelation factor);
PDBTitle: dimerization of gelation factor from dictyostelium2 discoideum: crystal structure of rod domains 5 and 6
98 c3n6yA_
not modelled
65.1
18
PDB header: unknown functionChain: A: PDB Molecule: immunoglobulin-like protein;
PDBTitle: crystal structure of an immunoglobulin-like protein (pa1606) from2 pseudomonas aeruginosa at 1.50 a resolution
99 c2kutA_
not modelled
63.3
11
PDB header: structural genomics, unknown functionChain: A: PDB Molecule: uncharacterized protein;
PDBTitle: solution structure of gmr58a from geobacter metallireducens.2 northeast structural genomics consortium target gmr58a
100 c3ogrA_
not modelled
62.1
20
PDB header: hydrolaseChain: A: PDB Molecule: beta-galactosidase;
PDBTitle: complex structure of beta-galactosidase from trichoderma reesei with2 galactose
101 c1xc6A_
not modelled
62.0
18
PDB header: hydrolaseChain: A: PDB Molecule: beta-galactosidase;
PDBTitle: native structure of beta-galactosidase from penicillium sp. in complex2 with galactose
102 c3qe5A_
not modelled
60.5
14
PDB header: cell adhesionChain: A: PDB Molecule: major cell-surface adhesin pac;
PDBTitle: complete structure of streptococcus mutans antigen i/ii carboxy-2 terminus
103 d2dj4a1
not modelled
60.1
19
Fold: Immunoglobulin-like beta-sandwichSuperfamily: E set domainsFamily: Filamin repeat (rod domain)
104 d1vjja1
not modelled
59.8
6
Fold: Immunoglobulin-like beta-sandwichSuperfamily: E set domainsFamily: Transglutaminase N-terminal domain
105 c2ds4A_
not modelled
59.4
17
PDB header: protein bindingChain: A: PDB Molecule: tripartite motif protein 45;
PDBTitle: solution structure of the filamin domain from human2 tripartite motif protein 45
106 d1wlha1
not modelled
58.3
16
Fold: Immunoglobulin-like beta-sandwichSuperfamily: E set domainsFamily: Filamin repeat (rod domain)
107 c3pmmA_
not modelled
56.5
16
PDB header: hydrolaseChain: A: PDB Molecule: putative cytoplasmic protein;
PDBTitle: the crystal structure of a possible member of gh105 family from2 klebsiella pneumoniae subsp. pneumoniae mgh 78578
108 c2zxcA_
not modelled
54.5
11
PDB header: hydrolaseChain: A: PDB Molecule: neutral ceramidase;
PDBTitle: seramidase complexed with c2
109 d2d7na1
not modelled
54.3
13
Fold: Immunoglobulin-like beta-sandwichSuperfamily: E set domainsFamily: Filamin repeat (rod domain)
110 d2diaa1
not modelled
52.3
21
Fold: Immunoglobulin-like beta-sandwichSuperfamily: E set domainsFamily: Filamin repeat (rod domain)
111 c2x5pA_
not modelled
51.8
15
PDB header: protein bindingChain: A: PDB Molecule: fibronectin binding protein;
PDBTitle: crystal structure of the streptococcus pyogenes fibronectin binding2 protein fbab-b
112 c1f00I_
not modelled
49.2
10
PDB header: cell adhesionChain: I: PDB Molecule: intimin;
PDBTitle: crystal structure of c-terminal 282-residue fragment of2 enteropathogenic e. coli intimin
113 c2di7A_
not modelled
49.2
9
PDB header: structural proteinChain: A: PDB Molecule: bk158_1;
PDBTitle: solution structure of the filamin domain from human bk158_12 protein
114 d1g4ma2
not modelled
49.1
14
Fold: Immunoglobulin-like beta-sandwichSuperfamily: E set domainsFamily: Arrestin/Vps26-like
115 d2d7pa1
not modelled
48.9
18
Fold: Immunoglobulin-like beta-sandwichSuperfamily: E set domainsFamily: Filamin repeat (rod domain)
116 c2e6jA_
not modelled
48.0
15
PDB header: structural genomics, unknown functionChain: A: PDB Molecule: hydin protein;
PDBTitle: solution structure of the c-terminal papd-like domain from2 human hydin protein
117 d1ayra2
not modelled
47.4
9
Fold: Immunoglobulin-like beta-sandwichSuperfamily: E set domainsFamily: Arrestin/Vps26-like
118 c2okmA_
not modelled
46.1
11
PDB header: cell adhesionChain: A: PDB Molecule: collagen adhesin;
PDBTitle: crystal structure of ace19, the collagen binding subdomain2 of enterococcus faecalis surface protein ace
119 c2z1pA_
not modelled
45.4
12
PDB header: cell adhesionChain: A: PDB Molecule: collagen adhesin protein;
PDBTitle: the enterococcus faecalis mscramm ace binds its ligands by2 the collagen hug model
120 c3n9tA_
not modelled
45.0
14
PDB header: oxidoreductaseChain: A: PDB Molecule: pnpc;
PDBTitle: cryatal structure of hydroxyquinol 1,2-dioxygenase from pseudomonas2 putida dll-e4