Emaill.a.kelley@imperial.ac.uk
DescriptionP76578
DateWed Jan 25 15:21:09 GMT 2012
Unique Job IDbcbb97527c014f13

Detailed template information 

#Template Alignment Coverage3D Model Confidence% i.d. Template Information
1c2pn5A_



100.0 14 PDB header:immune system
Chain: A: PDB Molecule:thioester-containing protein i;
PDBTitle: crystal structure of tep1r
2c3cu7A_



100.0 14 PDB header:immune system
Chain: A: PDB Molecule:complement c5;
PDBTitle: human complement component 5
3c2b39B_



100.0 16 PDB header:immune system
Chain: B: PDB Molecule:c3;
PDBTitle: structure of mammalian c3 with an intact thioester at 3a resolution
4c2a73B_



100.0 15 PDB header:immune system
Chain: B: PDB Molecule:complement c3;
PDBTitle: human complement component c3
5c3g6jB_



100.0 15 PDB header:immune system
Chain: B: PDB Molecule:complement c3 alpha chain;
PDBTitle: c3b in complex with a c3b specific fab
6c3pvmB_



100.0 18 PDB header:immune system
Chain: B: PDB Molecule:cobra venom factor;
PDBTitle: structure of complement c5 in complex with cvf
7d1hzfa_



100.0 19 Fold:alpha/alpha toroid
Superfamily:Terpenoid cyclases/Protein prenyltransferases
Family:Complement components
8c1hzfA_



100.0 19 PDB header:immune system
Chain: A: PDB Molecule:complement factor c4a;
PDBTitle: c4adg fragment of human complement factor c4a
9d1c3da_



100.0 19 Fold:alpha/alpha toroid
Superfamily:Terpenoid cyclases/Protein prenyltransferases
Family:Complement components
10c3hrzA_



100.0 13 PDB header:immune system
Chain: A: PDB Molecule:cobra venom factor;
PDBTitle: cobra venom factor (cvf) in complex with human factor b
11d1qqfa_



100.0 16 Fold:alpha/alpha toroid
Superfamily:Terpenoid cyclases/Protein prenyltransferases
Family:Complement components
12c2qkiA_



100.0 14 PDB header:immune system/hydrolase inhibitor
Chain: A: PDB Molecule:complement c3;
PDBTitle: human c3c in complex with the inhibitor compstatin
13c3hs0B_



100.0 14 PDB header:immune system
Chain: B: PDB Molecule:cobra venom factor;
PDBTitle: cobra venom factor (cvf) in complex with human factor b
14c2a74B_



99.9 11 PDB header:immune system
Chain: B: PDB Molecule:complement component c3c;
PDBTitle: human complement component c3c
15c2p9rA_



99.7 23 PDB header:signaling protein
Chain: A: PDB Molecule:alpha-2-macroglobulin;
PDBTitle: human alpha2-macroglogulin is composed of multiple domains,2 as predicted by homology with complement component c3
16d2sqca1



99.6 16 Fold:alpha/alpha toroid
Superfamily:Terpenoid cyclases/Protein prenyltransferases
Family:Terpene synthases
17c3hrzC_



99.6 10 PDB header:immune system
Chain: C: PDB Molecule:cobra venom factor;
PDBTitle: cobra venom factor (cvf) in complex with human factor b
18c1o79A_



99.6 20 PDB header:isomerase
Chain: A: PDB Molecule:squalene--hopene cyclase;
PDBTitle: structures of human oxidosqualene cyclase inhibitors bound2 to a homolgous enzyme
19c1w6kA_



99.4 18 PDB header:isomerase
Chain: A: PDB Molecule:lanosterol synthase;
PDBTitle: structure of human osc in complex with lanosterol
20d1edya_



99.4 15 Fold:Common fold of diphtheria toxin/transcription factors/cytochrome f
Superfamily:Alpha-macroglobulin receptor domain
Family:Alpha-macroglobulin receptor domain
21d1ayoa_



not modelled 99.4 16 Fold:Common fold of diphtheria toxin/transcription factors/cytochrome f
Superfamily:Alpha-macroglobulin receptor domain
Family:Alpha-macroglobulin receptor domain
22d1bv8a_



not modelled 99.4 14 Fold:Common fold of diphtheria toxin/transcription factors/cytochrome f
Superfamily:Alpha-macroglobulin receptor domain
Family:Alpha-macroglobulin receptor domain
23d3es6b1



not modelled 99.4 11 Fold:Immunoglobulin-like beta-sandwich
Superfamily:E set domains
Family:SVA-like
24d2sqca2



not modelled 99.3 15 Fold:alpha/alpha toroid
Superfamily:Terpenoid cyclases/Protein prenyltransferases
Family:Terpene synthases
25d1w6ka1



not modelled 99.3 18 Fold:alpha/alpha toroid
Superfamily:Terpenoid cyclases/Protein prenyltransferases
Family:Terpene synthases
26c2qkiC_



not modelled 99.2 7 PDB header:immune system/hydrolase inhibitor
Chain: C: PDB Molecule:complement c3;
PDBTitle: human c3c in complex with the inhibitor compstatin
27d3dssb1



not modelled 98.6 17 Fold:alpha/alpha toroid
Superfamily:Terpenoid cyclases/Protein prenyltransferases
Family:Protein prenyltransferases
28c3e37B_



not modelled 98.2 13 PDB header:transferase
Chain: B: PDB Molecule:protein farnesyltransferase subunit beta;
PDBTitle: protein farnesyltransferase complexed with bisubstrate2 ethylenediamine scaffold inhibitor 5
29d2h6fb1



not modelled 98.1 13 Fold:alpha/alpha toroid
Superfamily:Terpenoid cyclases/Protein prenyltransferases
Family:Protein prenyltransferases
30c1cwvA_



not modelled 97.8 10 PDB header:structural protein
Chain: A: PDB Molecule:invasin;
PDBTitle: crystal structure of invasin: a bacterial integrin-binding protein
31c3q7aB_



not modelled 97.7 14 PDB header:transferase/transferase inhibitor
Chain: B: PDB Molecule:farnesyltransferase beta subunit;
PDBTitle: cryptococcus neoformans protein farnesyltransferase in complex with2 fpp and l-778,123
32c2pmvB_



not modelled 97.4 11 PDB header:transport protein
Chain: B: PDB Molecule:gastric intrinsic factor;
PDBTitle: crystal structure of human intrinsic factor- cobalamin complex at 2.62 a resolution
33c1tnuL_



not modelled 97.4 12 PDB header:transferase
Chain: L: PDB Molecule:geranylgeranyl transferase type i beta subunit;
PDBTitle: rat protein geranylgeranyltransferase type-i complexed with2 a ggpp analog and a gcincckvl peptide derived from rhob
34d1n4qb_



not modelled 97.4 12 Fold:alpha/alpha toroid
Superfamily:Terpenoid cyclases/Protein prenyltransferases
Family:Protein prenyltransferases
35d1w6ka2



not modelled 97.1 19 Fold:alpha/alpha toroid
Superfamily:Terpenoid cyclases/Protein prenyltransferases
Family:Terpene synthases
36c3draB_



not modelled 97.0 10 PDB header:transferase
Chain: B: PDB Molecule:geranylgeranyltransferase type i beta subunit;
PDBTitle: candida albicans protein geranylgeranyltransferase-i2 complexed with ggpp
37d1w8oa1



not modelled 96.8 17 Fold:Immunoglobulin-like beta-sandwich
Superfamily:E set domains
Family:E-set domains of sugar-utilizing enzymes
38c1l9mB_



not modelled 96.6 15 PDB header:transferase
Chain: B: PDB Molecule:protein-glutamine glutamyltransferase e3;
PDBTitle: three-dimensional structure of the human transglutaminase 32 enzyme: binding of calcium ions change structure for3 activation
39d1r76a_



not modelled 96.5 23 Fold:alpha/alpha toroid
Superfamily:Family 10 polysaccharide lyase
Family:Family 10 polysaccharide lyase
40d1gxma_



not modelled 96.5 23 Fold:alpha/alpha toroid
Superfamily:Family 10 polysaccharide lyase
Family:Family 10 polysaccharide lyase
41c2v3nA_



not modelled 96.1 13 PDB header:transport protein
Chain: A: PDB Molecule:transcobalamin-2;
PDBTitle: crystallographic analysis of upper axial ligand2 substitutions in cobalamin bound to transcobalamin
42d2afaa1



not modelled 95.1 17 Fold:alpha/alpha toroid
Superfamily:Six-hairpin glycosidases
Family:N-acylglucosamine (NAG) epimerase
43c2kl6A_



not modelled 94.6 19 PDB header:structural genomics, unknown function
Chain: A: PDB Molecule:uncharacterized protein;
PDBTitle: solution nmr structure of the cardb domain of pf1109 from2 pyrococcus furiosus. northeast structural genomics3 consortium target pfr193a
44c3e8vA_



not modelled 93.1 21 PDB header:structural genomics, unknown function
Chain: A: PDB Molecule:possible transglutaminase-family protein;
PDBTitle: crystal structure of a possible transglutaminase-family2 protein proteolytic fragment from bacteroides fragilis
45d1cwva1



not modelled 92.5 11 Fold:Immunoglobulin-like beta-sandwich
Superfamily:Invasin/intimin cell-adhesion fragments
Family:Invasin/intimin cell-adhesion fragments
46c1wlhA_



not modelled 91.7 12 PDB header:structural protein
Chain: A: PDB Molecule:gelation factor;
PDBTitle: molecular structure of the rod domain of dictyostelium2 filamin
47d1h8la1



not modelled 91.5 13 Fold:Prealbumin-like
Superfamily:Carboxypeptidase regulatory domain-like
Family:Carboxypeptidase regulatory domain
48c3irpX_



not modelled 90.5 36 PDB header:cell adhesion
Chain: X: PDB Molecule:uro-adherence factor a;
PDBTitle: crystal structure of functional region of uafa from staphylococcus2 saprophyticus at 1.50 angstrom resolution
49c3kptA_



not modelled 89.9 18 PDB header:cell adhesion
Chain: A: PDB Molecule:collagen adhesion protein;
PDBTitle: crystal structure of bcpa, the major pilin subunit of2 bacillus cereus
50c2h0eA_



not modelled 89.4 12 PDB header:hydrolase
Chain: A: PDB Molecule:transthyretin-like protein pucm;
PDBTitle: crystal structure of pucm in the absence of substrate
51d1qfha1



not modelled 89.3 18 Fold:Immunoglobulin-like beta-sandwich
Superfamily:E set domains
Family:Filamin repeat (rod domain)
52d1f00i1



not modelled 88.9 10 Fold:Immunoglobulin-like beta-sandwich
Superfamily:Invasin/intimin cell-adhesion fragments
Family:Invasin/intimin cell-adhesion fragments
53c1kv3F_



not modelled 88.8 9 PDB header:transferase
Chain: F: PDB Molecule:protein-glutamine gamma-glutamyltransferase;
PDBTitle: human tissue transglutaminase in gdp bound form
54c2l0dA_



not modelled 88.8 14 PDB header:cell adhesion
Chain: A: PDB Molecule:cell surface protein;
PDBTitle: solution nmr structure of putative cell surface protein ma_4588 (272-2 376 domain) from methanosarcina acetivorans, northeast structural3 genomics consortium target mvr254a
55c3qvaB_



not modelled 87.5 33 PDB header:hydrolase
Chain: B: PDB Molecule:transthyretin-like protein;
PDBTitle: structure of klebsiella pneumoniae 5-hydroxyisourate hydrolase
56c1g0dA_



not modelled 86.6 15 PDB header:transferase
Chain: A: PDB Molecule:protein-glutamine gamma-glutamyltransferase;
PDBTitle: crystal structure of red sea bream transglutaminase
57d1cwva2



not modelled 86.3 8 Fold:Immunoglobulin-like beta-sandwich
Superfamily:Invasin/intimin cell-adhesion fragments
Family:Invasin/intimin cell-adhesion fragments
58c2pz4A_



not modelled 85.2 16 PDB header:cell adhesion
Chain: A: PDB Molecule:protein gbs052;
PDBTitle: crystal structure of spab (gbs52), the minor pilin in gram-positive2 pathogen streptococcus agalactiae
59d1kgia_



not modelled 85.0 11 Fold:Prealbumin-like
Superfamily:Transthyretin (synonym: prealbumin)
Family:Transthyretin (synonym: prealbumin)
60d2q3za3



not modelled 84.8 8 Fold:Immunoglobulin-like beta-sandwich
Superfamily:Transglutaminase, two C-terminal domains
Family:Transglutaminase, two C-terminal domains
61c1h8lA_



not modelled 84.8 13 PDB header:carboxypeptidase
Chain: A: PDB Molecule:carboxypeptidase gp180 residues 503-882;
PDBTitle: duck carboxypeptidase d domain ii in complex with gemsa
62d1uwya1



not modelled 84.6 13 Fold:Prealbumin-like
Superfamily:Carboxypeptidase regulatory domain-like
Family:Carboxypeptidase regulatory domain
63d2diba1



not modelled 84.4 17 Fold:Immunoglobulin-like beta-sandwich
Superfamily:E set domains
Family:Filamin repeat (rod domain)
64c2r39A_



not modelled 83.9 17 PDB header:structural genomics, unknown function
Chain: A: PDB Molecule:fixg-related protein;
PDBTitle: crystal structure of fixg-related protein from vibrio parahaemolyticus
65c3mn8A_



not modelled 83.4 10 PDB header:hydrolase
Chain: A: PDB Molecule:lp15968p;
PDBTitle: structure of drosophila melanogaster carboxypeptidase d isoform 1b2 short
66c1uwyA_



not modelled 82.9 11 PDB header:hydrolase
Chain: A: PDB Molecule:carboxypeptidase m;
PDBTitle: crystal structure of human carboxypeptidase m
67c2h1xB_



not modelled 82.7 18 PDB header:hydrolase
Chain: B: PDB Molecule:5-hydroxyisourate hydrolase (formerly known as
PDBTitle: crystal structure of 5-hydroxyisourate hydrolase (formerly2 known as trp, transthyretin related protein)
68d1vjja3



not modelled 82.6 18 Fold:Immunoglobulin-like beta-sandwich
Superfamily:Transglutaminase, two C-terminal domains
Family:Transglutaminase, two C-terminal domains
69d1f86a_



not modelled 82.6 13 Fold:Prealbumin-like
Superfamily:Transthyretin (synonym: prealbumin)
Family:Transthyretin (synonym: prealbumin)
70d2e9ia1



not modelled 82.3 14 Fold:Immunoglobulin-like beta-sandwich
Superfamily:E set domains
Family:Filamin repeat (rod domain)
71c1f13A_



not modelled 81.7 10 PDB header:coagulation factor
Chain: A: PDB Molecule:cellular coagulation factor xiii zymogen;
PDBTitle: recombinant human cellular coagulation factor xiii
72c3saeA_



not modelled 81.7 9 PDB header:lyase
Chain: A: PDB Molecule:alpha-bisabolene synthase;
PDBTitle: structure of a three-domain sesquiterpene synthase: a prospective2 target for advanced biofuels production
73d1tfpa_



not modelled 81.3 16 Fold:Prealbumin-like
Superfamily:Transthyretin (synonym: prealbumin)
Family:Transthyretin (synonym: prealbumin)
74d1ttaa_



not modelled 80.8 11 Fold:Prealbumin-like
Superfamily:Transthyretin (synonym: prealbumin)
Family:Transthyretin (synonym: prealbumin)
75d1g0da3



not modelled 80.7 13 Fold:Immunoglobulin-like beta-sandwich
Superfamily:Transglutaminase, two C-terminal domains
Family:Transglutaminase, two C-terminal domains
76d1oo2a_



not modelled 80.6 18 Fold:Prealbumin-like
Superfamily:Transthyretin (synonym: prealbumin)
Family:Transthyretin (synonym: prealbumin)
77c2k7pA_



not modelled 80.4 14 PDB header:structural protein
Chain: A: PDB Molecule:filamin-a;
PDBTitle: filamin a ig-like domains 16-17
78c2gpzC_



not modelled 80.2 24 PDB header:hydrolase
Chain: C: PDB Molecule:transthyretin-like protein;
PDBTitle: transthyretin-like protein from salmonella dublin
79d1vjja2



not modelled 79.8 21 Fold:Immunoglobulin-like beta-sandwich
Superfamily:Transglutaminase, two C-terminal domains
Family:Transglutaminase, two C-terminal domains
80d1ex0a2



not modelled 77.5 11 Fold:Immunoglobulin-like beta-sandwich
Superfamily:Transglutaminase, two C-terminal domains
Family:Transglutaminase, two C-terminal domains
81d2q3za2



not modelled 77.4 12 Fold:Immunoglobulin-like beta-sandwich
Superfamily:Transglutaminase, two C-terminal domains
Family:Transglutaminase, two C-terminal domains
82d2d7ma1



not modelled 77.0 18 Fold:Immunoglobulin-like beta-sandwich
Superfamily:E set domains
Family:Filamin repeat (rod domain)
83d1qfha2



not modelled 76.6 10 Fold:Immunoglobulin-like beta-sandwich
Superfamily:E set domains
Family:Filamin repeat (rod domain)
84d1ex0a3



not modelled 76.2 17 Fold:Immunoglobulin-like beta-sandwich
Superfamily:Transglutaminase, two C-terminal domains
Family:Transglutaminase, two C-terminal domains
85c2xzzA_



not modelled 74.1 14 PDB header:transferase
Chain: A: PDB Molecule:protein-glutamine gamma-glutamyltransferase k;
PDBTitle: crystal structure of the human transglutaminase 1 beta-barrel domain
86c2nsmA_



not modelled 73.5 20 PDB header:hydrolase
Chain: A: PDB Molecule:carboxypeptidase n catalytic chain;
PDBTitle: crystal structure of the human carboxypeptidase n (kininase i)2 catalytic domain
87d2dmca1



not modelled 71.9 12 Fold:Immunoglobulin-like beta-sandwich
Superfamily:E set domains
Family:Filamin repeat (rod domain)
88c2ww8A_



not modelled 71.8 24 PDB header:cell adhesion
Chain: A: PDB Molecule:cell wall surface anchor family protein;
PDBTitle: structure of the pilus adhesin (rrga) from streptococcus2 pneumoniae
89c2e9jA_



not modelled 71.7 21 PDB header:structural protein
Chain: A: PDB Molecule:filamin-b;
PDBTitle: solution structure of the 14th filamin domain from human2 filamin-b
90d2dmba1



not modelled 70.9 25 Fold:Immunoglobulin-like beta-sandwich
Superfamily:E set domains
Family:Filamin repeat (rod domain)
91c2k7qA_



not modelled 69.3 13 PDB header:structural protein
Chain: A: PDB Molecule:filamin-a;
PDBTitle: filamin a ig-like domains 18-19
92d1cwva3



not modelled 67.9 8 Fold:Immunoglobulin-like beta-sandwich
Superfamily:Invasin/intimin cell-adhesion fragments
Family:Invasin/intimin cell-adhesion fragments
93d1v05a_



not modelled 67.4 17 Fold:Immunoglobulin-like beta-sandwich
Superfamily:E set domains
Family:Filamin repeat (rod domain)
94d1g0da2



not modelled 67.0 9 Fold:Immunoglobulin-like beta-sandwich
Superfamily:Transglutaminase, two C-terminal domains
Family:Transglutaminase, two C-terminal domains
95c3k11A_



not modelled 67.0 13 PDB header:hydrolase
Chain: A: PDB Molecule:putative glycosyl hydrolase;
PDBTitle: crystal structure of putative glycosyl hydrolase (np_813087.1) from2 bacteroides thetaiotaomicron vpi-5482 at 1.80 a resolution
96c1ksrA_



not modelled 65.8 15 PDB header:actin binding protein
Chain: A: PDB Molecule:gelation factor;
PDBTitle: the repeating segments of the f-actin cross-linking2 gelation factor (abp-120) have an immunoglobulin fold, nmr,3 20 structures
97c1qfhB_



not modelled 65.1 13 PDB header:actin binding protein
Chain: B: PDB Molecule:protein (gelation factor);
PDBTitle: dimerization of gelation factor from dictyostelium2 discoideum: crystal structure of rod domains 5 and 6
98c3n6yA_



not modelled 65.1 18 PDB header:unknown function
Chain: A: PDB Molecule:immunoglobulin-like protein;
PDBTitle: crystal structure of an immunoglobulin-like protein (pa1606) from2 pseudomonas aeruginosa at 1.50 a resolution
99c2kutA_



not modelled 63.3 11 PDB header:structural genomics, unknown function
Chain: A: PDB Molecule:uncharacterized protein;
PDBTitle: solution structure of gmr58a from geobacter metallireducens.2 northeast structural genomics consortium target gmr58a
100c3ogrA_



not modelled 62.1 20 PDB header:hydrolase
Chain: A: PDB Molecule:beta-galactosidase;
PDBTitle: complex structure of beta-galactosidase from trichoderma reesei with2 galactose
101c1xc6A_



not modelled 62.0 18 PDB header:hydrolase
Chain: A: PDB Molecule:beta-galactosidase;
PDBTitle: native structure of beta-galactosidase from penicillium sp. in complex2 with galactose
102c3qe5A_



not modelled 60.5 14 PDB header:cell adhesion
Chain: A: PDB Molecule:major cell-surface adhesin pac;
PDBTitle: complete structure of streptococcus mutans antigen i/ii carboxy-2 terminus
103d2dj4a1



not modelled 60.1 19 Fold:Immunoglobulin-like beta-sandwich
Superfamily:E set domains
Family:Filamin repeat (rod domain)
104d1vjja1



not modelled 59.8 6 Fold:Immunoglobulin-like beta-sandwich
Superfamily:E set domains
Family:Transglutaminase N-terminal domain
105c2ds4A_



not modelled 59.4 17 PDB header:protein binding
Chain: A: PDB Molecule:tripartite motif protein 45;
PDBTitle: solution structure of the filamin domain from human2 tripartite motif protein 45
106d1wlha1



not modelled 58.3 16 Fold:Immunoglobulin-like beta-sandwich
Superfamily:E set domains
Family:Filamin repeat (rod domain)
107c3pmmA_



not modelled 56.5 16 PDB header:hydrolase
Chain: A: PDB Molecule:putative cytoplasmic protein;
PDBTitle: the crystal structure of a possible member of gh105 family from2 klebsiella pneumoniae subsp. pneumoniae mgh 78578
108c2zxcA_



not modelled 54.5 11 PDB header:hydrolase
Chain: A: PDB Molecule:neutral ceramidase;
PDBTitle: seramidase complexed with c2
109d2d7na1



not modelled 54.3 13 Fold:Immunoglobulin-like beta-sandwich
Superfamily:E set domains
Family:Filamin repeat (rod domain)
110d2diaa1



not modelled 52.3 21 Fold:Immunoglobulin-like beta-sandwich
Superfamily:E set domains
Family:Filamin repeat (rod domain)
111c2x5pA_



not modelled 51.8 15 PDB header:protein binding
Chain: A: PDB Molecule:fibronectin binding protein;
PDBTitle: crystal structure of the streptococcus pyogenes fibronectin binding2 protein fbab-b
112c1f00I_



not modelled 49.2 10 PDB header:cell adhesion
Chain: I: PDB Molecule:intimin;
PDBTitle: crystal structure of c-terminal 282-residue fragment of2 enteropathogenic e. coli intimin
113c2di7A_



not modelled 49.2 9 PDB header:structural protein
Chain: A: PDB Molecule:bk158_1;
PDBTitle: solution structure of the filamin domain from human bk158_12 protein
114d1g4ma2



not modelled 49.1 14 Fold:Immunoglobulin-like beta-sandwich
Superfamily:E set domains
Family:Arrestin/Vps26-like
115d2d7pa1



not modelled 48.9 18 Fold:Immunoglobulin-like beta-sandwich
Superfamily:E set domains
Family:Filamin repeat (rod domain)
116c2e6jA_



not modelled 48.0 15 PDB header:structural genomics, unknown function
Chain: A: PDB Molecule:hydin protein;
PDBTitle: solution structure of the c-terminal papd-like domain from2 human hydin protein
117d1ayra2



not modelled 47.4 9 Fold:Immunoglobulin-like beta-sandwich
Superfamily:E set domains
Family:Arrestin/Vps26-like
118c2okmA_



not modelled 46.1 11 PDB header:cell adhesion
Chain: A: PDB Molecule:collagen adhesin;
PDBTitle: crystal structure of ace19, the collagen binding subdomain2 of enterococcus faecalis surface protein ace
119c2z1pA_



not modelled 45.4 12 PDB header:cell adhesion
Chain: A: PDB Molecule:collagen adhesin protein;
PDBTitle: the enterococcus faecalis mscramm ace binds its ligands by2 the collagen hug model
120c3n9tA_



not modelled 45.0 14 PDB header:oxidoreductase
Chain: A: PDB Molecule:pnpc;
PDBTitle: cryatal structure of hydroxyquinol 1,2-dioxygenase from pseudomonas2 putida dll-e4